Chester Cricket needs help. That's the message John Robin carries into the Times Square subway station where Harry Cat and Tucker Mouse live. Quickly, Chester's good friends set off on the long, hard journey to the Old Meadow, where all is not well.

Houses are creeping closer. Bulldozers and construction are everywhere. It looks like Chester and his friends' home will be ruined and the children of the town won't have a place to play. Harry Cat and Tucker Mouse are used to the city life. Now in the country, they need to find a place to stay and good things to eat. And most of all they must think of a plan to help their friends.